明朝年間，錦衣衛陸欽因罪被發配充軍，途中因緣際會，與樊英押送貪官張嵩回京指認通敵太監王公公。經過生死追逃，最終陸欽犧牲，張嵩抵京，卻被王公公處死于監牢。多年后，王公公被樊英擊殺，善惡終得報。

唐載初年，位于郊外校場倒塌的通天浮屠殘骸一夜之間復活。天空中一條火龍照亮夜空，并噴射火球摧毀了整個校場，死傷無數。隨后通天浮屠也燃起大火，化為灰燼。而異族鬼道在城外叫囂，只有武則天效仿大佛自焚，天下才可免受災難。城中內外人心惶惶，狄仁杰奉命徹查此案，卻遇到了與自己有過節的大理寺少卿裴東升。二人不打不相識，被迫一起調查此案。面對外敵的入侵，隊友的不信任，狄仁杰決心背水一戰，勢要查明案件的真相。

　　After breaking out of a rehabilitation zoo, Frankie, a squirrel with all the angles, returns to his forest. He learns from his dim-witted best friend, Cody, a porcupine and street-con, that while he was in lock-up, the entire forest was picked clean by The A.C.O.R.N.S. Institute (Alternative Cosmetic Oils for Regenerative Natural Skin-care). No acorn was left behind. With winter just around the corner, it's a potentially devastating blow to all forest critters. So, Frankie has no choice but to bust into the brand new, state-of-the-art, rodent-proof A.C.O.R.N.S. Processing Plant and take back all of the acorns! Though a seemingly impossible heist, success would not only save the lives of his forest friends, but it would also give Frankie a chance to win back the love of his life, Lola, a local "show squirrel." Sadly, Lola has been burnt by too many of Frankie's false promises in the past. With the support of Frankie's old mentor, Mr. Bellwood, a wealthy badger, Frankie and Cody assemble a team of top notch "crackers" from all over the forest, including: a bat named Fly Boy, a top notch munitions expert; Raitch, a computer hacking albino snake, who never allows her lack of appendages to get in the way; Edsy a tiny "family frog," whose ability to freeze himself solid for days on end without harm is invaluable; a germophobic French rat named Liam, who can collapse his skeleton to get into any space; Chinese father-daughter pangolins named Pango and Lin, the best diggers in the forest; and Bellwood's hedgehog butler, Oscar who knows how to get his hands on anything the crew needs. With winter knocking at their door, Frankie and his crew have only three days to prepare and execute this unprecedented heist. When they learn that the processing of their forest's acorns has been pushed up a day, Lola's refusal to participate leaves Frankie and his crew one critter short. With everything on the line, Frankie and his crew of Animal Crackerz set out to pull off the biggest heist in animal history!

當兼顧工作和撫養孩子的單身母親成熙（尹世雅飾）為她年幼的女兒在整個房子里安裝家庭攝像頭時，奇怪的事情開始了。