媒體從業人員帕特里克·卡特（肯塔克爾·德利 Kentucker Audley 飾）意外聯系到了失蹤許久的姐姐卡羅琳（艾米·西米茨 Amy Seimetz 飾）。卡羅琳當初曾在戒毒所治療，如今住進了一個名為“圣伊甸園”的地方，并自稱過上了烏托邦般的生活。為了弄清姐姐的下落，帕特里克和同事兼好友山姆·特納（AJ·鮑恩 AJ Bowen 飾）、杰克·威廉姆斯（喬·斯萬博格 Joe Swanberg 飾）啟程前往圣伊甸園的所在地調查。這座樂園位于一個鮮為人知的隱秘所在，許許多多生活中經歷痛苦和坎坷的人們，在“父親”（吉恩·瓊斯 Gene Jones 飾）的感召和撫慰下重獲新生。　　出于媒體人的敏感，山姆取得了和“父親”對談的機會，但他卻從對方幽默機智的談吐中嗅到了一絲殺氣……

　　After breaking out of a rehabilitation zoo, Frankie, a squirrel with all the angles, returns to his forest. He learns from his dim-witted best friend, Cody, a porcupine and street-con, that while he was in lock-up, the entire forest was picked clean by The A.C.O.R.N.S. Institute (Alternative Cosmetic Oils for Regenerative Natural Skin-care). No acorn was left behind. With winter just around the corner, it's a potentially devastating blow to all forest critters. So, Frankie has no choice but to bust into the brand new, state-of-the-art, rodent-proof A.C.O.R.N.S. Processing Plant and take back all of the acorns! Though a seemingly impossible heist, success would not only save the lives of his forest friends, but it would also give Frankie a chance to win back the love of his life, Lola, a local "show squirrel." Sadly, Lola has been burnt by too many of Frankie's false promises in the past. With the support of Frankie's old mentor, Mr. Bellwood, a wealthy badger, Frankie and Cody assemble a team of top notch "crackers" from all over the forest, including: a bat named Fly Boy, a top notch munitions expert; Raitch, a computer hacking albino snake, who never allows her lack of appendages to get in the way; Edsy a tiny "family frog," whose ability to freeze himself solid for days on end without harm is invaluable; a germophobic French rat named Liam, who can collapse his skeleton to get into any space; Chinese father-daughter pangolins named Pango and Lin, the best diggers in the forest; and Bellwood's hedgehog butler, Oscar who knows how to get his hands on anything the crew needs. With winter knocking at their door, Frankie and his crew have only three days to prepare and execute this unprecedented heist. When they learn that the processing of their forest's acorns has been pushed up a day, Lola's refusal to participate leaves Frankie and his crew one critter short. With everything on the line, Frankie and his crew of Animal Crackerz set out to pull off the biggest heist in animal history!

 影片開始于兩個都叫瑪麗的少女（Jitka Cerhová 和 Ivana Karbanová 飾）的對話：沒有人理解我們，這個世界上的一切正在變得越來越糟糕。于是兩人決定：我們也要變得越來越壞！瑪麗一號喜歡和老男人約會，在他們吃飯的時候，瑪麗二號會突然出現，旁若無人地大吃特吃。瑪麗一號把老男人送上火車，老男人以為她會陪著他，火車開動，她卻下了火車。瑪麗二號在一個收藏蝴蝶標本的男青年家中，用他的標本遮住自己身體的隱秘部位。她們在夜總會里搗亂，最后被趕走。她們無意中闖入一個沒有人的宴會，開始肆無忌憚地享受美食.....本片獻給精神生活一團糟的人們。